I would take the R310 offline and try rebooting all 4 R320 from the POE switch.
The highest firmware that the R310 supports is 184.108.40.206.121 (you can't add it or unless you downgrade all the R320 to 220.127.116.11.121 and then you can add r310. However that means you are stuck at that firmware)
Ruckus hasn't pushed 18.104.22.168.233 on their websites yet... for some reason only selected APs got 22.214.171.124.233 through unleashed online upgrade.... I haven't bothered to upgrade yet and stuck to 212 for til they released it on the websites. I would rather wait since sees there's some kernel panic issues..